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Another class of wall proteins, like collagen, contain high levels of hydroxyproline. These proteins arethought to strengthen the wall, and they are produced in greatly increasedamounts as a local responseto attack by pathogens. From the genome sequenceof Arabidopsis,it has been estimated that more than 700 genes are required tosynthesize, assemble,and remodel the plant cell wall.
Some of the main polymers found in the primary and secondary cell wall are listed in Table l9-g.pflmarycellwallprasmam e mo r a n e50 nmFigure19-79Scalemodel of a portionof a primaryplant cellwall showingthetwo major polysaccharidenetworks.Theorthogonallyarrangedlayersof cellulosemicrofibrils(green)aretied into a networkby the cross-linkingglycans(red)thatform hydrogenbondswith themicrofibrils.This network is coextensivewith a networkor matrixof pecticpolysaccharides(blud. fhe networkofcelluloseand cross-linkingglycansprovidestensilestrength,whilethepectinnetworkresistscompression.Cellulose,cross-linkingglycans,anopectinaretypicallypresentin roughlyequalamountsin a primarycellwall.Themiddlelamellais especiallyrichin pectin,and it cementsadjacentcellstogether.())ttt(8)('s].raqoULlila)pue sllaMupuBJo{sapnot'y) 'puedxaue) slla)luauoduro)s]! r])uM ut uotl)altpaql Iq UPdur paurrlralapsr'looqse s eq ) n s ' u e b r ou e ; o a d e q sl e u 4a q f'leuateuJlleMMauJo uotlrasutaql qllM]ra)uo) ur srn)lo uorsuedxallal'ql6uallsa;rsua11ear6a^eqq)rqM'sluqUorlr,laq] Jo uorleluauoJo la^ellsotuJauura q ] o t r e l n ) r p u a d r audo r l > a r repu l I ; 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